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    But note that some authors place Barbus grypus in the Indian genus Tor and that evidently more work needs to be done on its relationships and on those of other species that have no evident Euro-Mediterranean relatives, but whose origins may well lie in the Oriental Region.Other Middle Eastern cyprinid genera are regarded by Durand et al.Cyprinion may have entered the Middle East during the colonization event that isolated the genera Barbus sensu lato and Schizothorax in the European and Asian basins respectively.The divergence of these species is similar in time to the radiation of the Leuciscinae supposedly centred in Siberia based on fossil records. (sic) from the Lower Miocene of Saudi Arabia showing an early date for the entry of cyprinids to the Afro-Arabian Plate.

    The first pectoral and the first pelvic fin ray are unbranched and not included in counts.The Middle East leuciscine cyprinids have Europe as an important Palearctic influence consistent with the Lago Mare dispersion while the the cyprinine cyprinids show three highly divergent lineages, namely one shared with the Euro-Mediterranean area (Barbus/Luciobarbus), a relict of the Lago Mare dispersion, one shared with Africa (Carasobarbus/Varicorhinus subgenus) and one with Asia (Garra).The Lago Mare dispersion occurred during a salinity crisis in the Mediterranean Sea 5.5 MY ago in the Late Miocene when freshwater fish were able to disperse through oligohaline or fresh water in the Paratethys Sea to reach the Middle East (Bianco, 1990). (2002) conflict with this scenario - the Carasobarbus clade that includes Barbus (= Tor) grypus shows a separation divergence later than the salinity crisis in the Pliocene when no migration route was available.Pharyngeal teeth lie on a modified, fifth gill arch which can be seen or probed behind the shoulder girdle, just inside the gill opening.The arch has to be removed with dissecting equipment to count the teeth.

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